Denise Applewhite/Office of Communications
IN MEMORIAM: Biology professor emeritus JOHN BONNER died Feb. 7 in Portland, Ore. He was 98. Bonner joined the faculty in 1947, serving three times as chair of the biology department. After becoming emeritus in 1990, he continued his research for more than two decades at Princeton and in Oregon.

Bonner devoted most of his life to studying cellular slime molds. He published 20 books on their behavior as well as on broader themes in evolutionary biology — the last of which, Randomness in Evolution, was published when he was 93. He received an honorary degree from Princeton in 2006.

Friends of the Princeton University Library
IN MEMORIAM: BEN PRIMER, former associate university librarian for rare books and special collections, died Feb. 11 in Princeton after a long illness. He worked in the University Library for 25 years, beginning in 1990 as the curator of public-policy papers and then for a decade as University archivist, expanding access to Mudd Library’s holdings and widening its collections. He served as associate university librarian from 2002 to 2014, playing a role in planning the Firestone Library renovations and invigorating the Friends of the Princeton University Library.